It gets better.. so, I plugged my WiGig card into an ASRock H270M Pro4 board, and no matter where I put that card, the NVidia graphics card slot remained at x16. I had it in Slot 1, and the WiGig in Slot 4 as per https://www.asrock.com/MB/Intel/H270M%20Pro4/index.asp#Specification and set it to PCIE 3.0 in the UEFI. Either GPU-Z isn't correctly reading the slot speed, which should have dropped to x8, or the UEFI is smart enough to recognize the Intel WiGig card as not being a graphics card, and routes it as PCM traffic instead.
So add another variable to the list - motherboard provider or version. Gorrammit. Either way, the WiGig card isn't going to perform any better on my systems no matter what slot I put it in.